Bonnerichthys

Bonnerichthys Temporal range: Late Cretaceous, 89–66 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Restoration
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	†Pachycormiformes
Family:	†Pachycormidae
Genus:	†Bonnerichthys Friedman et al., 2010
Species:	†B. gladius
Binomial name
†Bonnerichthys gladius (Cope, 1873)
Synonyms
Portheus gladius Cope, 1873 Protosphyraena gladius (Cope, 1873)

Bonnerichthys is a genus of fossil fishes within the family Pachycormidae that lived during the Coniacian to Maastrichtian stage of the Late Cretaceous.^[1] Fossil remains of this taxon were first described from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas (Late Coniacian-Early Campanian), and additional material was later reported from the Pierre Shale, Mooreville Chalk, Demopolis Chalk, Wenonah Formation, and Moreno Formation, among other localities.^[2] It grew to at least 5 metres (16 ft) in total body length,^[1] substantially less than the related Leedsichthys from the Jurassic which likely grew up to 16.5 metres (54 ft).^[3]

One of the most significant features of Bonnerichthys is the recognition that it was a filter feeder, living on plankton. This recognition that many large-bodied fish from the Mesozoic in the Pachycormidae were filter feeders shows that this niche was filled for at least 100 million years before previously known. The modern niche is filled by several species of sharks and the baleen whales.

The international team that described the genus named it after the Marion Charles Bonner fossil hunting family,^[4] whose collections from the Niobrara Cretaceous chalk of western Kansas are in many museums and research institutions.

• Bonnerichthys at the Oceans of Kansas website. Includes detailed taxonomic history, life restorations, bibliography, many photos of fossil remains.
• Bonnerichthys discussion at boneblogger.com

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Teleostei sensu de Pinna, 1996 (= Teleosteomorpha)

Kingdom: Animalia Phylum: Chordata Class: Actinopterygii Subclass: Neopterygii
Actinopterygii see Actinopterygii Neopterygii see Neopterygii Teleostei see below↓

Teleostei
Teleostei †Atacamichthys †Baleiichthys †Barschichthys †Bunoderma †Ceramurus †Dorsetichthys †Elpistoichthys †Eurycormus †Galkinia †Grimmenichthys †Hengnania †Hulettia? †Hungkiichthys †Ichthyokentema †Jialingichthys †Koonwarria †Ligulella †Luxembourgichthys †Majokia †Marcopoloichthys †Neobaleiichthys †Oligopleurus? †Pholidophorichthys †Pholidophoroides †Pholidophoropsis †Prohalecites †Ruedersdorfia †Seefeldia †Seinstedtia †Signeuxella †Pachycormidae Asthenocormus Australopachycormus Bonnerichthys Eugnathides? Euthynotus Germanostomus Haasichthys Hypsocormus Kaykay Leedsichthys Martillichthys Notodectes Ohmdenia Orthocormus Pachycormus Protosphyraena Pseudoasthenocormus Rhinconichthys Sauropsis Saurostomus Simocormus †Aspidorhynchidae Aspidorhynchus Belonostomus Jonoichthys Pseudovinctifer Richmondichthys Vinctifer †Pholidophoridae sensu stricto Annaichthys Ceneichthys Eopholidophorus Knerichthys Lombardichthys Malingichthys Parapholidophorus Pholidoctenus Pholidophoretes Pholidophorus Pholidorhynchodon Pseudopholidoctenus Zambellichthys †Archaeomaenidae Aphnelepis Archaeomaene Gurvanichthys Oreochima Wadeichthys Zaxilepis †Siyuichthyidae Bogdaichthys Dsungarichthys Manasichthys Siyuichthys Wukangia †Pleuropholidae Austropleuropholis Gondwanapleuropholis Parapleuropholis Pleuropholis Zurupleuropholis †Ankylophoridae Ankylophorus Lehmanophorus Neopholidophoropsis Pholidophoristion Siemensichthys Songaichthys Steurbautichthys †Catervariolidae Catervariolus Kisanganichthys Songanella Teleosts with cycloid scales see below↓ Otolith taxa †Archengraulis †Bicavolithus †Doggerichthys †Sphaeronchus †Xenoleptolepis

Teleosts with cycloid scales
Teleosts with cycloid scales †Antofagastaichthys †Araripichthys †Paraclupavus †Pholidolepis †Todiltia †Leptolepidae Leptolepis Longileptolepis Proleptolepis †Luisiellidae Cavenderichthys? Luisiella Waldmanichthys? †Ascalaboidae Ascalabos Ebertichthys Tharsis †Crossognathiformes Bavarichthys Chongichthys Salminops? Varasichthyidae? Bobbichthys Domeykos Luisichthys Protoclupea Varasichthys Notelopidae Bavarichthys? Michin? Notelops Nusaviichthys Crossognathidae Apsopelix Crossognathus Pachyrhizodontidae Apricenapiscis Aquilopiscis Cavinichthys Chongichthys? Elopopsis Goulmimichthys Greenwoodella Kradimus Lebrunichthys Michin? Motlayoichthys Nardopiscis Pachyrhizodus Platinx Polcynichthys Rhacolepis Stanhopeichthys Tingitanichthys †Ichthyodectiformes Africathrissops Allothrissops Altamuraichthys Antarctithrissops Ascalabothrissops? Capassoichthys Carsothrissops Dugaldia Faugichthys Furloichthys Garganoichthys Gillicus Occithrissops Ogunichthys Pachythrissops? Prymnetes Sultanuvaisia Thrissops Verraesichthys Chuhsiungichthyidae Chuhsiungichthys Jinjuichthys Mesoclupea Bardackichthyidae Amakusaichthys? Bardackichthys Heckelichthys? Cladocyclidae Aidachar Chirocentrites Chiromystus Cladocyclus Cladocynodon Eubiodectes Ichthyodectidae Cooyoo Ghrisichthys Ichthyodectes Postredectes Xiphactinus Saurodontidae Gwawinapterus Prosaurodon Saurocephalus Saurodon Unamichthys Vallecillichthys Crown-Teleostei see below↓

Crown-Teleostei
Elopomorpha see Elopomorpha †Orthogonikleithridae Cavenderichthys? Leptolepides Orthogonikleithrus Waldmanichthys? Osteoglossomorpha see Osteoglossomorpha †Tselfatiiformes Eoplethodus Protobramidae Abisaadichthys Eusebichthys Protobrama Plethodidae Bachea Bananogmius Concavotectum Dixonanogmius Enischnorhynchus Luxilites Martinichthys Moorevillia Niobrara Paranogmius Pentanogmius Plethodus Pseudonogmius Pseudothryptodus Rhamphoichthys Syntegmodus Thryptodus Tselfatia Zanclites Clupeocephala Otocephala Euteleostei

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