Leptusa

Leptusa
Leptusa carolinensis
Nomenklatur
Leptusa Kraatz, 1856
Hører til
småkortvinger, Kortvinger, Polyphaga, biller, insekter
Økologi
Antall arter:		> 400, ca. 260 i Europa, 4 arter funnet i Norge
Utbredelse:		alle verdensdeler bortsett fra Australia og Madagaskar
Inndelt i

Leptusa er en slekt av biller som hører til den store underfamilien småkortvinger (Aleocharinae) i familien kortvinger (Staphylinidae). Det er beskrevet mer enn 400 arter, fire av dem er funnet i Norge.

Utseende

Forholdsvis små (gjerne 2-4 millimeter lange), middels slanke kortvinger, kroppen nokså jevnbred. På farge er de ofte rødlige eller brunlige. Hodet er ganske bredt med nokså små fasettøyne. Pronotum er bredere enn langt, med rundede sider og hjørner. Dekkvingene er forholdsvis korte og har bakkanten tydelig innbuktet innenfor bakhjørnene. Føttene er fire-leddete.

Levevis

Leptusa-artene er vanlige på skogbunnen, og kan også finnes under løs bark på døde trær.

Systematisk inndeling / norske arter

Denne artsrike slekten blir delt opp i ca. 80 underslekter.

Treliste

Ordenen biller, Coleoptera
- Underordenen Polyphaga
  - Overfamilien åtsel- og rovbiller, Staphylinoidea
    - Familien kortvinger (Staphylinidae) Latreille, 1802
      - Underfamilien småkortvinger (Aleocharinae) Fleming, 1821
        Stammen Homalotini Heer, 1839
        Slekten Leptusa Kraatz, 1856
        Leptusa pulchella (Mannerheim, 1831)
        
        Leptusa fumida (Erichson, 1839)
        
        Leptusa norvegica A. Strand, 1941
        
        Leptusa ruficollis (Erichson, 1839)

Liste over artene

Leptusa abchasica Bernhauer, 1936
Leptusa abdominalis (Motschulsky, 1851)
Leptusa abeillei Bernhauer, 1900
Leptusa acuta Assing, 2002
Leptusa aisenicola Pace, 2001
Leptusa albergianensis Pace, 1989
Leptusa allobrogica Pace, 2000
Leptusa alpicola Brancsik, 1874
Leptusa amisensis Pace, 1982
Leptusa andina (Fauvel, 1866)
Leptusa angustiarumberninae Scheerpeltz, 1972
Leptusa anmashanensis Pace, 1996
Leptusa annapurnensis Pace, 1982
Leptusa antarctica (Kiesenwetter, 1877)
Leptusa apennina Holdhaus, 1924
Leptusa apfelbeckiana Likovský, 1984
Leptusa araucana Pace, 2001
Leptusa areraensis Scheerpeltz, 1973
Leptusa argentierensis Pace, 1999
Leptusa armatissima Assing, 2008
Leptusa armeniaca Pace, 1989
Leptusa artviniensis Pace, 1982
Leptusa asiatica Bernhauer, 1909
Leptusa asperata Eppelsheim, 1888
Leptusa assingi Pace, 1996
Leptusa asturica Pace, 1983
Leptusa asturiensis Eppelsheim, 1880
Leptusa athosensis Zerche, 2002
Leptusa atriceps (Waterhouse, 1875)
Leptusa atrocephala Bernhauer, 1905
Leptusa austriaca Scheerpeltz, 1935
Leptusa baldensis Ganglbauer, 1895
Leptusa baldomontis Scheerpeltz, 1972
Leptusa bellinorum Assing, 2004
Leptusa benacensis Pace, 1980
Leptusa bengalensis Pace, 1982
Leptusa bergamasca Scheerpeltz, 1972
Leptusa bernhaueri Scheerpeltz, 1926
Leptusa binaghii Pace, 1979
Leptusa biumbonata Scheerpeltz, 1973
Leptusa blesioi Pace, 1981
Leptusa bonvouloirii Brisout de Barneville, 1867
Leptusa bozdaghensis Pace, 1983
Leptusa brachati Pace, 1980
Leptusa brevicollis Casey, 1893
Leptusa brigantii Pace, 1979
Leptusa brixiensis Pace, 1979
Leptusa brucki W. Scriba, 1867
Leptusa cabrerensis Pace, 1981
Leptusa cadiensis Pace, 1989
Leptusa calancensis Szallies, 2014
Leptusa californiana Pace, 1989
Leptusa calliceroides Assing, 2004
Leptusa camunnensis Pace, 1981
Leptusa canonica Casey, 1906
Leptusa cansigliensis Pace, 1975
Leptusa cantabrica Paśnik, 1998
Leptusa carinata Bernhauer, 1909
Leptusa carniolica Pace, 1983
Leptusa carolinensis Pace, 1989
Leptusa carpathica J. Weise, 1877
Leptusa casalei Pace, 1979
Leptusa cassagnaui Pace, 1984
Leptusa caucasica Eppelsheim, 1878
Leptusa cautinensis Pace, 2001
Leptusa cavallensis Pace, 1980
Leptusa cavallicola Pace, 1980
Leptusa cavata Assing, 2003
Leptusa centralis Pace, 1992
Leptusa cephalotes Bernhauer, 1909
Leptusa ceresoleana Pace, 1979
Leptusa cerrutii Coiffait, 1970
Leptusa chengduensis Pace, 2001
Leptusa chilensis (Fauvel, 1866)
Leptusa chinensis Pace, 1997
Leptusa cimmeria Pace, 1996
Leptusa circassica Bernhauer, 1935
Leptusa coiffaiti Pace, 1989
Leptusa confinis Pace, 1982
Leptusa cordicollis Portevin, 1929
Leptusa cornigera Assing, 2002
Leptusa corsica Pace, 1989
Leptusa crassiventris Scheerpeltz, 1973
Leptusa cremata Assing, 2003
Leptusa crenulata Bernhauer, 1899
Leptusa cribrata Pace, 1996
Leptusa cribratula (Casey, 1906)
Leptusa cribripennis Kraatz, 1856
Leptusa crinita Assing, 2007
Leptusa cultellata Assing, 2008
Leptusa cuneensis Pace, 1979
Leptusa curvata Assing, 2006
Leptusa cypria Pace, 1996
Leptusa delphiniformis Assing, 2009
Leptusa deplanata (K. Sawada, 1970)
Leptusa deprehendens Pace, 1981
Leptusa devoluyiensis Pace, 1989
Leptusa diecki Pace, 1983
Leptusa difficilis Pace, 1996
Leptusa difformis (Mulsant & Rey, 1853)
Leptusa discolor Assing, 2006
Leptusa dissimilans Assing, 2004
Leptusa dubia
Leptusa durmitorensis Pace, 1984
Leptusa elegans
Leptusa emplenotoides Assing, 2006
Leptusa epirensis Pace, 1989
Leptusa erlangensis Pace, 1999
Leptusa estrelensis Pace, 1981
Leptusa etrusca Pace, 1979
Leptusa europaea Pace, 1996
Leptusa everestensis Pace, 1989
Leptusa excaecata Assing, 2002
Leptusa eximia
Leptusa fauciumberninae Scheerpeltz, 1973
Leptusa fauciumredortae Scheerpeltz, 1972
Leptusa fauveli Eppelsheim, 1889
Leptusa ferroi
Leptusa fibula Assing, 2003
Leptusa filiformis Scheerpeltz, 1972
Leptusa fischeri Bernhauer, 1910
Leptusa flagellata Assing, 2002
Leptusa flagrifera Assing, 2009
Leptusa flavicornis Brancsik, 1874
Leptusa florae Scheerpeltz, 1926
Leptusa formidabilis Pace, 1996
Leptusa foroiuliensis Pace, 1980
Leptusa franzi Pace, 1981
Leptusa franziana Pace, 1981
Leptusa frontalis
Leptusa fuliginosa
Leptusa fumida
Leptusa gadesensis Pace, 1982
Leptusa gaisbergeri
Leptusa gansuensis Pace, 1997
Leptusa ganzica Assing, 2002
Leptusa gatineauensis
Leptusa gerlitzensis Pace, 1983
Leptusa ghoropanensis Pace, 1989
Leptusa gibbera Assing, 2011
Leptusa gimmeli
Leptusa glabriceps Bernhauer, 1923
Leptusa globulicollis (Mulsant & Rey, 1853)
Leptusa gonggamontis Pace, 1997
Leptusa gracilenta Pace, 1983
Leptusa gracilipes
Leptusa gracilis
Leptusa gracillima Pace, 1983
Leptusa granulicauda Eppelsheim, 1890
Leptusa granulipennis Eppelsheim, 1880
Leptusa grignaensis Scheerpeltz, 1973
Leptusa hamulata Assing, 2010
Leptusa hastata Assing, 2002
Leptusa helambuica Assing, 2009
Leptusa hercegovinensis Bernhauer, 1910
Leptusa herminia Pace, 1982
Leptusa hiarens Assing, 2011
Leptusa himalayiana Pace, 1989
Leptusa hlavaci Assing, 2006
Leptusa hlisnikovskyi Pace, 1983
Leptusa hoelzeli Scheerpeltz, 1948
Leptusa honshuica Assing, 2002
Leptusa hopffgarteni Eppelsheim, 1881
Leptusa hummleri Bernhauer, 1910
Leptusa hummleriana Bernhauer, 1935
Leptusa ilamensis Pace, 1987
Leptusa ilsae Bernhauer, 1935
Leptusa improvisa Assing, 2009
Leptusa indica
Leptusa inexpectata
Leptusa ionopolitana Pace, 1982
Leptusa ishizuchiensis Pace, 1982
Leptusa italica Pace, 1977
Leptusa janbellini Assing, 2001
Leptusa janczyki Pace, 1983
Leptusa janetscheki Scheerpeltz, 1956
Leptusa japonica
Leptusa jeanneli Scheerpeltz, 1935
Leptusa jelineki Pace, 1983
Leptusa jinfomontis Pace, 2001
Leptusa jiudingensis Pace, 1999
Leptusa jucunda
Leptusa juliana Pace, 1980
Leptusa jurassica
Leptusa kahleni Pace, 1980
Leptusa kangdingensis Pace, 2010
Leptusa kaohsiungensis Pace, 2007
Leptusa kappenbergeri
Leptusa karawankarum Pace, 1983
Leptusa kaszabi Pace, 1981
Leptusa kaufmanni
Leptusa khnzoriani Pace, 1982
Leptusa kitazawai (K. Sawada, 1970)
Leptusa knabli Bernhauer, 1909
Leptusa knappeorum Scheerpeltz, 1964
Leptusa kocae Bernhauer, 1905
Leptusa kochiana Bernhauer, 1936
Leptusa komareki Pace, 1983
Leptusa koralpicola Pace, 1983
Leptusa korgei Scheerpeltz, 1970
Leptusa koronensis
Leptusa kosmajensis Pace, 1987
Leptusa lackneri Assing, 2011
Leptusa laevicauda (Scheerpeltz, 1958)
Leptusa laeviuscula
Leptusa lapidicola
Leptusa laticeps Scheerpeltz, 1973
Leptusa lativentris
Leptusa lederi Eppelsheim, 1883
Leptusa leonhardi
Leptusa leonica Pace, 1981
Leptusa leptotyphloides
Leptusa lessiniensis Pace, 1978
Leptusa limata Assing, 2002
Leptusa linderi Scheerpeltz, 1966
Leptusa linderiana Scheerpeltz, 1966
Leptusa loebli Pace, 1984
Leptusa lohsei Pace, 1989
Leptusa lombarda Bernhauer, 1905
Leptusa longalata Assing, 2017
Leptusa longilobata Assing, 2007
Leptusa lophophororum Pace, 1987
Leptusa luigionii Bernhauer, 1935
Leptusa luremontis Pace, 1999
Leptusa lurensis Pace, 1989
Leptusa magellanica Pace, 1989
Leptusa majevicensis Scheerpeltz, 1935
Leptusa major Bernhauer, 1900
Leptusa maleschevskaensis
Leptusa manasluensis Assing, 2002
Leptusa mancinii Pace, 1980
Leptusa mandli Scheerpeltz, 1953
Leptusa manfredi Pace, 1980
Leptusa marasica Assing, 2006
Leptusa marianii Pace, 1979
Leptusa marieluiseae
Leptusa marmotae Assing, 2002
Leptusa martensi Pace, 1987
Leptusa media Scheerpeltz, 1973
Leptusa merditana
Leptusa merkli Bernhauer, 1900
Leptusa meschniggi Bernhauer, 1935
Leptusa meybohmi Assing, 2014
Leptusa michai Assing, 2002
Leptusa microphthalma
Leptusa microvolans Pace, 1997
Leptusa migrituber Assing, 2017
Leptusa minutissima Pace, 1983
Leptusa moczarskii Bernhauer, 1915
Leptusa monacha (Fauvel, 1898)
Leptusa monachorum Bernhauer, 1912
Leptusa monguzzii Pace, 1989
Leptusa montana (Fauvel, 1866)
Leptusa montiscathedrae Pace, 1978
Leptusa montisgrappae Pace, 1975
Leptusa montisgrignae Scheerpeltz, 1973
Leptusa montisparii Scheerpeltz, 1953
Leptusa montispasubii Pace, 1975
Leptusa montiumcarnorum (Scheerpeltz, 1958)
Leptusa mucronemontis Scheerpeltz, 1972
Leptusa munelensis
Leptusa muscicola
Leptusa myrmidon
Leptusa nahuelbutana
Leptusa nanula
Leptusa nemoricultrix Pace, 1995
Leptusa nepalica Scheerpeltz, 1976
Leptusa nepalorum Pace, 1982
Leptusa netolitzkyi Bernhauer, 1914
Leptusa newar Pace, 1989
Leptusa nigerrima Pace, 1983
Leptusa nitidiventris
Leptusa nonveilleri Pace, 1984
Leptusa norica Pace, 1979
Leptusa norvegica
Leptusa notabilis Scheerpeltz, 1972
Leptusa nubigena
Leptusa numidica Pace, 1983
Leptusa nurdaghensis Assing, 2003
Leptusa obirensis Scheerpeltz, 1935
Leptusa obscura
Leptusa occulta Pace, 1975
Leptusa ochsi Pace, 1983
Leptusa oertzeni Eppelsheim, 1888
Leptusa olympica Pace, 1994
Leptusa opaca
Leptusa opacina Pace, 2001
Leptusa oreophila
Leptusa oropaensis Pace, 1979
Leptusa orumboviorum Pace, 1982
Leptusa osellai Pace, 1977
Leptusa pangeoensis Assing & Wunderle, 1999
Leptusa paradoxa Pace, 1980
Leptusa parallela
Leptusa parvibulbata Assing, 2008
Leptusa pascuorum Pace, 1975
Leptusa pasubiana Scheerpeltz, 1953
Leptusa pedemontana Bernhauer, 1935
Leptusa peinantamontis Pace, 2007
Leptusa pelionensis Bernhauer, 1916
Leptusa peninsulae Pace, 1989
Leptusa peregrina Pace, 1995
Leptusa peristerica Pace, 1989
Leptusa persica Assing, 2009
Leptusa petzeniensis Pace, 1989
Leptusa peyerimhoffi Bernhauer, 1935
Leptusa piceata (Mulsant & Rey, 1853)
Leptusa piciformis Assing, 2009
Leptusa pilatensis Scheerpeltz, 1972
Leptusa pingtungensis Pace, 1995
Leptusa pinkeri
Leptusa pirinensis
Leptusa pizzocolensis Pace, 1981
Leptusa plitvicensis Pace, 1980
Leptusa poggii Pace, 1979
Leptusa pollicita Assing, 2010
Leptusa portusnaoniensis Pace, 1989
Leptusa pratensis Pace, 1989
Leptusa priesneri Pace, 1989
Leptusa proiecta Assing, 2008
Leptusa pseudoalpestris Scheerpeltz, 1935
Leptusa pseudocaucasica Pace, 1983
Leptusa pseudopaca
Leptusa pseudosmokyiensis
Leptusa puellaris
Leptusa puetzi Assing, 2008
Leptusa pulchella
Leptusa punctithorax Bernhauer, 1935
Leptusa punctulata (Mulsant & Rey, 1875)
Leptusa pusio
Leptusa puthzi Assing, 2002
Leptusa pyrenaica Pace, 1982
Leptusa qinlingensis Pace, 1999
Leptusa quinqueimpressa Assing, 2008
Leptusa rambouseki
Leptusa recta Assing, 2006
Leptusa reinosensis Pace, 1996
Leptusa reitteri Eppelsheim, 1880
Leptusa repentina Scheerpeltz, 1972
Leptusa rhaetoromanica
Leptusa rhilensis Pace, 1983
Leptusa rizensis Pace, 1996
Leptusa rorata Pace, 1995
Leptusa rosai Pace, 1979
Leptusa roscidavallensis Pace, 1981
Leptusa rossica Bernhauer, 1915
Leptusa rougemonti Pace, 1997
Leptusa rousi Pace, 1983
Leptusa ruffoi Pace, 1983
Leptusa ruficollis
Leptusa ruficornis
Leptusa rugatipennis
Leptusa sabauda Pace, 1989
Leptusa salernitana Pace, 1979
Leptusa salonichia Bernhauer, 1914
Leptusa samia Assing, 2004
Leptusa sampeyrensis Pace, 2009
Leptusa sanfilippoi Pace, 1980
Leptusa sarensis Pace, 1983
Leptusa savonensis Pace, 1977
Leptusa scabripennis (Mulsant & Rey, 1875)
Leptusa schaschli
Leptusa schawalleri Assing, 2011
Leptusa schoenmanni Pace, 1983
Leptusa schuelkei Pace, 1999
Leptusa secreta Bernhauer, 1900
Leptusa semiscabra Scheerpeltz, 1972
Leptusa semivolans Pace, 1996
Leptusa sengleti Pace, 1984
Leptusa serbica Scheerpeltz, 1935
Leptusa seriana Scheerpeltz, 1973
Leptusa serullazi
Leptusa shaanxiensis Pace, 1999
Leptusa sibirica Pace, 1982
Leptusa sibyllinica Pace, 1977
Leptusa sica Assing, 2003
Leptusa sichuanensis Pace, 1997
Leptusa simoni Eppelsheim, 1878
Leptusa sinorum Pace, 2001
Leptusa smetanai Pace, 1989
Leptusa smetanaiella Pace, 1989
Leptusa smetanaorum Pace, 1989
Leptusa smokyiensis Pace, 1989
Leptusa soganlica Assing, 2007
Leptusa solieri Pace, 1989
Leptusa soriciformis Assing, 2009
Leptusa sororella Pace, 1989
Leptusa sparsa (Fauvel, 1898)
Leptusa spirarum Pace, 1996
Leptusa spoliata Assing, 2002
Leptusa stimulans Assing, 2008
Leptusa stocksi
Leptusa stoeckleini Bernhauer, 1914
Leptusa storkani Roubal, 1917
Leptusa subalpina
Leptusa subcarpathica Roubal, 1930
Leptusa subcaucasica Pace, 1983
Leptusa subconvexa (Mulsant & Rey, 1875)
Leptusa subnivalis (Roubal, 1911)
Leptusa substricta Assing, 2017
Leptusa sudetica Lokay, 1900
Leptusa sulcicollis Bernhauer, 1932
Leptusa svanetica Assing, 2017
Leptusa taichii Kishimoto, 2008
Leptusa taichungensis Pace, 1996
Leptusa taiwanensis Pace, 1992
Leptusa tarvisiana Pace, 1989
Leptusa taurica Assing, 2004
Leptusa taurinensis Pace, 1989
Leptusa tectusoides Assing, 2002
Leptusa temperei Pace, 1989
Leptusa tenchiensis Pace, 1996
Leptusa tendamontis Scheerpeltz, 1972
Leptusa tendicola Pace, 1999
Leptusa tenerrima Bernhauer, 1900
Leptusa tenuicornis Assing, 2006
Leptusa tenuissima Scheerpeltz, 1972
Leptusa thailandensis Pace, 2004
Leptusa tirolensis Bernhauer, 1900
Leptusa titillans Assing, 2002
Leptusa tomorensis Pace, 1989
Leptusa toumayeffi Scheerpeltz, 1972
Leptusa transversiceps Pace, 1983
Leptusa trapezuntis Pace, 1989
Leptusa triangulata Assing, 2017
Leptusa tricolor W. Scriba, 1870
Leptusa tridentina Pace, 1980
Leptusa trumplinensis Pace, 1981
Leptusa turgida Assing, 2006
Leptusa tusculanensis Pace, 1981
Leptusa ultracollensis Pace, 1982
Leptusa umbhakica Assing, 2011
Leptusa vailatii Pace, 1989
Leptusa vallisvenyi Scheerpeltz, 1972
Leptusa veniorum Pace, 1999
Leptusa ventosensis Pace, 1996
Leptusa venusta (Hochhuth, 1849)
Leptusa vercorensis Pace, 1989
Leptusa vermiensis Pace, 1981
Leptusa veronensis Pace, 1983
Leptusa vietnamensis Pace, 1992
Leptusa wechseliensis Pace, 1980
Leptusa weiratheri Scheerpeltz, 1972
Leptusa winkleri Scheerpeltz, 1926
Leptusa winkleriana Scheerpeltz, 1965
Leptusa winneguthi Linke, 1938
Leptusa winneguthiana Pace, 1983
Leptusa woerndlei Scheerpeltz, 1935
Leptusa wolongensis Assing, 2002
Leptusa wunderlei Pace, 1996
Leptusa wuyica Assing, 2002
Leptusa xanthopyga Eppelsheim, 1881
Leptusa xiahensis Pace, 1997
Leptusa xianensis Pace, 1999
Leptusa xuemontis Pace, 2001
Leptusa yakorum Pace, 1987
Leptusa yamato (K. Sawada, 1990)
Leptusa yunnanensis Pace, 2001
Leptusa zanettiorum Pace, 1980
Leptusa zerchei Pace, 1989
Leptusa zhemomontis Assing, 2010
Leptusa zhongdianensis Pace, 2010
Leptusa zoiai Pace, 1982

Kilder

Park, J.-S., C.E. Carlton og M.L. Ferro 2010. Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U.S.A., with descriptions of four new species. Zootaxa 2662: 1-27. [1]
Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki.
Fauna Europaea Web Service Version 2.4 (2011) Utbredelsesdatabase for europeiske dyr. [2] Arkivert 2. juni 2017 hos Wayback Machine.